Dragonflies and Damselflies: References

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Reference Link. Citation Primary biogeographic zone Disciplinary domains URL Usage
May 1992c. May, ML (1992) Morphological and Ecological Differences Among Species of Ladona (Anisoptera: Libellulidae). Bulletin of American Odonatology 1(3): 51-56 Nearctic Taxonomy Behaviour Ecology Link Check use

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Abstract
[Introduction] The three species placed by Needham (1897) in his new genus Ladona have suffered a checkered nomenclatorial history. Disagreement still exists as to 1) whether the taxon should be considered a full genus, a subgenus, or merely a synonym of Libellula; and 2) whether the included species are all truly distinct. In this note I use Ladona as a genus name for convenience, but I intend to take no particular position on the first question. If Kennedy's (1922) phylogeny of Libellula is accepted, then recognizing Ladona as a genus can only be justified if Neotetrum and Holotania, at least, are also accepted at that rank, but, on the other hand, that conclusion depends critically on Kennedy's rather problematic placement of L. quadrimaculata. ... At various times, both L. julia (Hagen) and L. deplanata (Rambur) have been synonymized with the first described (but probably least known) species, L. exusta (Say). Bennefield (1965) reviewed the taxonomic status of the group. He concluded that the three species are indeed distinct, but this view has not been universally accepted (e.g., Dunkle, 1989). In the present paper I confirm the validity of all three species, principally on the basis of penile morphology, and present preliminary evidence for a partial behavioral and ecological separation between L. exusta and L. deplanata where they co-occur in the New Jersey Pine Barrens.